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The thinking was that the temperature differential between the used upper portion and the unused (at least on our air cooled trannys) can cause moisture to form in the bottom reservoir, that when unused just "sits" there potentially leading to corrosion.
It's prolly not a big concern, but I didn't want to go down that road.
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You are correct. Temp changes will cause air-flow and condensation. Usually, they used to come with fittings to block the cooler off if sold as a multi-use.